Continuously updated Google Scholar Profile for AM Helton can be accessed here.
(Italicized names are postdoctoral associates, Underlined names are graduate students, and underlined and italicized names are undergraduate students who participated in the research within the publication while they were a part of Helton’s lab. The first author is corresponding unless indicated by *)
  1. Cummins, CS, AD Rosemond, NJ Tomczyk, SJ Wenger, PM Bumpers, V Gulis, AM Helton, JP Benstead. Consumer group and leaf species alter the temperature dependence of leaf litter breakdown.  Ecology. Accepted.
  2. Hare, DK, AM Helton, CS Cummins, PM Bumpers, NJ Tomczyk, PA Rogers, SJ Wenger, ER Hotchkiss, AD Rosemond, JP Benstead. 2024. Leaf litter breakdown phenology in headwater stream networks is modulated by groundwater thermal inputs and litter type. Limnology and Oceanography-Letters. https://doi.org/10.1002/lol2.10423
  3. Klionsky, S, C Neill, AM Helton, BA Lawrence. 2024. Groundwater seeps are hotspots for nitrogen removal in a restored cranberry bog. Biogeochemistry. https://doi.org/10.1007/s10533-024-01156-w
  4. Jackson, KE, EM Moore, AM Helton*, AB Haynes, JR Barclay, MA Briggs. 2024. Exploring landscape and geologic controls on spatial patterning of streambank groundwater discharge in a mixed land use watershed. Hydrologic Processes. https://doi.org/10.1002/hyp.15112
  5. O'Donnell, K, ES Bernhardt, X Yang, R Emanuel, M Ardón, M Lerdau, A Manda, A Braswell, TK BenDor, EC Edwards, E Frankenberg, AM Helton, JS Kominoski, A Lesen, L Naylor, G Noe, K Tully, E White, J Wright. 2024. Saltwater Intrusion and Sea Level Rise threatens coastal landscapes and communities. Anthropocene. https://doi.org/10.1016/j.ancene.2024.100427
  6. Moore, EM, JR Barclay, AB Haynes, KE Jackson, AM Bisson, MA Briggs, AM Helton. 2023. Where the past meets the present: Connecting nitrogen from watersheds to their streams through groundwater flow paths. Environmental Research Letters. 18 124039. DOI 10.1088/1748-9326/ad0c86
  7. Barry, A, SK Ooi, AM Helton, B Steven, CS Elphick, BA Lawrence*. 2023. Carbon dynamics vary among tidal marsh plant species in a sea-level rise experiment. Wetlands 43 (7) 78
  8. Haynes, AB, MA Briggs*, E Moore, K Jackson, J Knighton, DM Rey, AM Helton. 2023. Shallow and local or deep and regional? Inferring source groundwater characteristics across mainstem river discharge faces. Hydrological Processes 27 (7) e14939.
  9. Snarski, JW, M Dietz, AM Helton, J Knighton. 2023. Potential hydrologic pathways of deicing salt chloride transport evaluate with SWMM. Journal of Hydrologic Engineering. 28 (8) 04023022
  10. Hare, DK, SA Benz, BL Kurylyk, ZC Johnson, NC Terry, AM Helton. 2023. Paired air and stream temperature analysis (PASTA) to evaluate groundwater influence on streams. Water Resources Research. doi: 10.1029/2022WR033
  11. Helton, AM, JL Morse, EB Sudduth, M Ardón, R Bier, KA Voss, MRV Ross, JR Blaszczak, JE Brandt, M Simonin, JD Rocca, A Carter, JR Gerson, EA Ury, MJ Vlah. 2023. At the interfaces of the hydrologic sciences: Connecting water, elements, ecosystems, and people through the major contributions of Dr. Emily Bernhardt. Journal of Hydrology 619: 129251. doi: 10.1016/j.jhydrol.2023.129251
  12. Blaszczak, JR, LE Koenig, FH Mejia, L Gomez-Gener, CL Dutton, AM Carter, NB Grimm, JW Harvey, AM Helton, MJ Cohen. 2022. Global extent, patterns, and drivers of hypoxia in streams and rivers.  Limnology and Oceanography-Letters. doi: 10.1002/lol2.10297
  13. Swails, EE, M Ardon, KW Krauss, AL Peralta, RE Emmanuel, AM Helton, JL Morse, L Gutenberg, N Cormier, D. Shoch, S Settelmyer, E Soderholm, B Boutin, C Peoples, S Ward. 2022.  Response of soil respiration to changes in soil temperature and water table level in drained and restored peatlands of the southeastern United States. Carbon Balance and Management 17, 18. doi: 10.1186/s13021-022-00219-5
  14. Tomczyk, N, AD Rosemond, J Kominoski, D Manning, J Benstead, V Gulis, S Thomas, E Hotchkiss, AM Helton. 2022. Nitrogen and phosphorus uptake stoichiometry tracks supply ratio during two-year whole-ecosystem nutrient additions. Ecosystems. doi: 10.1007/s10021-022-00813-1
  15. Ooi, SK, A Barry, BA Lawrence, CS Elphick, AM Helton. 2022. Vegetation zones are indicators of denitrification potential in salt marshes. Ecological Applications. 32 (6) e2630
  16. Bernhardt, ES, P Savoy, MJ Vlah, AA Appling, LE Koenig, RO Hall, M Arriota, JR Blaszczak, AM Carter, M Cohen, NB Grimm, JW Harvey, JB Heffernan, AM Helton, JD Hosen, L Kirk, WH McDowell, JS Reed, EH Stanley, EG Stets, CB Yackulic.  2022. Light and flow regimes regulate the metabolism of rivers. Proceedings of the National Academy of Sciences. 119 (8) e2121976119
  17. Wollheim, WM. TK Harms, AL Robinson, LE Koenig, AM Helton C Song, WB Bowden, JC Finlay.  2022. Superlinear scaling of riverine biogeochemical function with watershed size. Nature Communications 13, 1230. doi: 10.1038/s41467-022-28630-z
  18. Barclay, JR, MA Briggs, JJ Starn, EM Moore, AEH Hanson, AM Helton. 2022. Where groundwater seeps: Evaluating modeled groundwater discharge patterns with thermal infrared surveys at the river network scale.  Advances in Water Research 160 (104108). https://doi.org/10.1016/j.advwatres.2021.104108
  19. Briggs, MA, KE Jackson, F Liu, EM Moore, A Bisson, AM Helton. 2022. Exploring local riverbank sediment controls on the occurrence of preferential groundwater discharge points. Water 14(1), 11. https://doi.org/10.3390/w14010011
  20. Rodriquez-Cardona, B, A Wymore, A Argerich, R Barnes, S Bernal, J Brookshire, A Coble, WK Dodds, H Fazekas, AM Helton, P Johnes, S Johnson, J Jones, S Kaushal, P Kortelainen, C Lopez-Lloreda, R Spences, W McDowell.  2021. Shifting Stoichiometry: Long-term trends in stream dissolved organic matter alter C:N ratios due to history of atmospheric acid deposition. Global Change Biology 28(1): 98-114. https://doi.org/10.1111/gcb.15965
  21. Wymore, AS, PJ Johnes, S Bernal, ENJ Brookshire, HM Fazekas, AM Helton, A Argerich, RT Barnes, AA Coble, WK Dodds, S Haq, SL Johnson, JB Jones, SS Kaushal, P Kortelainen, C Lopez-Lloreda, B Rodriquez-Cardona, RGM Spencer, PL Sullivan, CA Yates, WH McDowell. 2021. Gradients of Anthropogenic Nutrient Enrichment Alter N Composition and DOM Stoichiometry in Freshwater Ecosystems.  Global Biogeochemical Cycles. Doi: 10.1029/2021GB006953
  22. Walker, S, G Robbins, AM Helton, BA Lawrence. 2021. Road salt inputs alter biogeochemistry but not community composition in exurban forested wetlands. Ecosphere. https://doi.org/10.1002/ecs2.3814
  23. Barry, A, SK Ooi, AM Helton, S Blaire, C Elphick, BA Lawrence. 2021. Vegetation zonation predicts soil carbon mineralization and microbial communities in southern New England salt marshes. Estuaries and Coasts. https://doi.org/10.1007/s12237-021-00943-0
  24. Sullivan, CJ, JC Vokoun, AM Helton, MA Briggs, BL Kurylyk. 2021. An ecohydrological typology for thermal refuges in streams and rivers. Ecohydrology. https://doi.org/10.1002/eco.2295
  25. Hare, DK, AM Helton, ZC Johnson, JW Lane, MA Briggs. 2021. Continental-scale analysis of shallow and deep groundwater contributions to streams. Nature-Communications 12 (1450). doi: 10.1038/s41467-021-21651-0
  26. Granville, KE, SK Ooi, LE Koenig, BA Lawrence, CS Elphick, AM Helton. 2021. Seasonal patterns of denitrification and N2O production in a southern New England salt marsh. Wetlands 41(7) https://doi.org/10.1007/s13157-021-01393-x
  27. Barclay, JR, JJ Starn, MA Briggs, AM Helton. 2020. Improved prediction of management-relevant groundwater discharge characteristics throughout river networks. Water Resources Research. doi:10.1029/2020WR028027
  28. Macklem, DC, AM Helton, MW Tingley, JM Dickson, TAG Rittenhouse. 2020. Stream salamander persistence influenced by the interaction between exurban housing age and development. Urban Ecosystems.  23 (117): 117-132.
  29. Harvey, MC, DK Hare, A Hackman, G Davenport, AB Haynes, AM Helton, JW Lane, MA Briggs. 2019. Evaluation of stream and wetland restoration using UAS-based thermal infrared mapping. Water. 11(8), 1668, https://doi.org/10.3390/w11081568
  30. Koenig, LE, AM Helton, P Savoy, E Bertuzzo, JB Heffernan, RO Hall, ES Bernhardt. 2019. Emergent productivity regimes of river networks. Limnology and Oceanography-Letters. https://doi.org/10.1002/lol2.10115
  31. Schoepfer, VA, AJ Burgin, TD Loecke, AM Helton. 2019. Seasonal salinization decreases spatial heterogeneity of sulfate reducing activity. Soil Systems. 3(2), 25; https://doi.org/10.3390/soilsystems3020025
  32. Doroski, AA, AM Helton and TM Vadas. 2019. Denitrification Potential and Carbon Mineralization in Restored and Unrestored Coastal Wetland Soils Across an Urban Landscape. Wetlands. https://doi.org/10.1007/s13157-019-01128-z
  33. ​Helton, AM, M Ardon, ES Bernhardt. 2019. Hydrologic context alters greenhouse gas feedbacks of coastal wetland salinization. Ecosystems. https://doi.org/10.1007/s10021-018-0325-2
  34. Doroski, AA, AM Helton, T Vadas. 2019. Greenhouse gas fluxes from wetlands at the intersection of urban pollution and saltwater intrusion: a soil core experiment. Soil Biology and Biochemistry. 131: 44-53. https://doi.org/10.1016/j.soilbio.2018.12.023
  35. Reinhold, AM, GC Poole, C Izurieta, AM Helton, and ES Bernhardt.  2019. From conceptualization to computer code: Automatic generation of constraint-based models of biogeochemical systems.  Ecological Informatics.
  36. Ardon, M, AM Helton, ES Bernhardt. 2018. Salinity effects on greenhouse gas emissions from wetland soils are contingent upon hydrologic setting: a microcosm experiment. Biogeochemistry. 140:217-232. https://doi.org/10.1007/s10533-018-0486-2
  37. Song, C, WK Dodds, J Ruegg, A Argerich, CL Baker, WB Bowden, MM Douglas, KJ Farrell, MB Flinn, EA Garcia, AM Helton, TK Harms, S Jia, JB Jones, LE Koenig, JS Kominoski, WH McDowell, D McMaster, SP Parker, AD Rosemond, CM Ruffing, KR Sheehan, MT Trentman, MR Whiles, WM Wollheim, F Ballantyne. 2018. Continental-scale decrease in net primary productivity in streams due to climate warming. Nature Geosciences. doi: 10.1038/s41561-018-0125-5
  38. Carmichael, MJ, AM Helton, JC White, WK Smith. 2017. Standing dead trees are a conduit for the atmospheric flux of CH4 and CO2 from wetlands. Wetlands. 38: 133-143. https://doi.org/10.1007/s13157-017-0963-8
  39. Bertuzzo, E, AM Helton, RO Hall, TJ Battin. 2017. Scaling of dissolved organic carbon removal in river networks. Advances in Water Resources. 110: 136-146.
  40. Helton, AM, RO Hall, E Bertuzzo. 2017. How network structure can affect nitrogen removal by streams. Freshwater Biology. doi: 10.1111/fwb.12990 **Top 20 downloaded article June 2016 to July 2018.
  41. Ardon, M, AM Helton, MD Scheuerell, ES Bernhardt. 2017. Fertilizer legacies meet saltwater incursion: Challenges and constraints for coastal plain wetland restoration. Elementa: Science of the Anthropocene. Special Collection: Ghosts of land-use past: Do land-use legacy effects constrain the restoration of aquatic ecosystems? doi: http://doi.org/10.1525/elementa.236
  42. Barclay, J, H Tripp, CJ Bellucci, GS Warner, AM Helton. 2016. Do water body classifications predict water quality? Journal of Environmental Management. 183(1): 1-12.
  43. McInerney, E and AM Helton. 2016. The effects of soil moisture and vegetation on carbon emissions from constructed wetlands. Wetlands. 36: 275-284. doi:10.1007/s13157-016-0736-9.
  44. Ardon, M, AM Helton, ES Bernhardt. 2016. Drought and saltwater incursion synergistically reduce dissolved organic carbon export from coastal freshwater wetlands. Biogeochemistry. 127 (2-3): 411-426
  45. Helton, AM, M Ardon, ES Bernhardt. 2015. Thermodynamic constraints on the utility of ecological stoichiometry for explaining global biogeochemical patterns. Ecology Letters. 18 (10): 1049-1056. **Recommended by Faculty of 1000
  46. Tant, CJ, AD Rosemond, AM Helton, MR First. 2015. Nutrient enrichment alters the relative contribution of fungi, bacteria, and detritivores to leaf litter breakdown. Freshwater Science. 34 (4): 1259-1271.
  47. Helton, AM, MS Wright, ES Bernhardt, GC Poole, RM Cory, JA Stanford. 2015. Dissolved organic carbon lability increases with water residence time in the alluvial aquifer of a river floodplain ecosystem. Journal of Geophysical Research - Biogeosciences. 120. doi:10.1002/2014JG002832.
  48. Payn, RA, AM Helton, GC Poole, C Izurieta, ES Bernhardt, and AJ Burgin. 2014. A generalized model of aquatic microbial metabolism based on thermodynamic, kinetic, and stoichiometric theory. Ecological Modelling. 294: 1-18.
  49. Helton, AM, ES Bernhardt, A Fedders. 2014. Biogeochemical regime shifts in coastal landscapes: The contrasting effects of saltwater incursion and agricultural pollution on greenhouse gas emissions from a freshwater wetland. Biogeochemistry. 120: 133-147. doi: 10.1007/s10533-014-9986-x
  50. Hopfensperger, KN, AJ Burgin, VA Schoepfer, and AM Helton. 2014. Impacts of saltwater incursion on plant communities, anaerobic microbial metabolism, and resulting relationships in a restored freshwater wetland. Ecosystems. 17(5): 792-807. doi: 10.1007/s10021-014-9760-x.
  51. LINX collaborators: WK Dodds, JR Webster, CL Crenshaw, AM Helton, JM O'Brien, E Martí, AE Hershey, JL Tank, AJ Burgin, NB Grimm, SK Hamilton, DJ Sobota, GC Poole, JJ Beaulieu, LT Johnson, LR Ashkenas, RO Hall, Jr., SL Johnson, WM Wollheim, WB Bowden. 2014. The Lotic Intersite Nitrogen Experiments: an example of successful ecological research collaboration. Freshwater Science. 33(3):700–710. doi: 10.1086/676938
  52. Helton, AM, GC Poole, RA Payn, C Izurieta, and JA Stanford. 2014. Relative influences of the river channel, floodplain surface, and alluvial aquifer on simulated hydrologic residence time in a montane river floodplain. Geomorphology. Special Issue: “Discontinuities in Fluvial Systems”205:17-26. doi: 10.1016/j.geomorph.2012.01.004
  53. Mehring, AS,  RR Lowrance, AM Helton, G Vellidis, CM Pringle, and DD Bosch. 2013. Inter-annual drought length governs dissolved organic carbon dynamics in blackwater rivers of the western upper Suwannee River basin. Journal of Geophysical Research - Biogeosciences. 118: 1636–1645. doi: 10.1002/2013JG002415.
  54. Potter, JD, WH McDowell, ML Daley, and AM Helton. 2013. Incorporating urban infrastructure into biogeochemical assessment of urban tropical streams in Puerto Rico.  Biogeochemistry. doi: 10.1007/s10533-013-9914-5
  55. Luhr, R, D Reimanis, R Cross, C Izurieta, GC Poole, AM Helton. 2013. Natural Science Visualization Using Digital Theater Software: Adapting existing planetarium software to model ecological systems. Proceedings of the International Conference on Information Science and Applications. doi: 10.1109/ICISA.2013.6579381.
  56. Helton, AM, GC Poole, RA Payn, C Izurieta, JA Stanford. 2012. Scaling flow path processes to fluvial landscapes: An integrated field and model assessment of temperature and dissolved oxygen dynamics in a river-floodplain-aquifer system. Journal of Geophysical Research - Biogeosciences, Special Issue "Linking physical, chemical, and biological processes in watersheds from the cellular and grain scales to the landscape scale" 117, G00N14. doi:10.1029/2012JG002025
  57. Bernhardt, ES, BD Lutz, RS King, JP Fay, CE Carter, AM Helton, D Campagna, and J Amos.  2012. How many mountains can we mine? Assessing the regional degradation of Central Appalachian rivers by surface coal mining.  Environmental Science & Technology. 46 (15): 8115–8122. doi: 10.1021/es301144q
  58. Izurieta, C, GC Poole, RA Payn, AM Helton, I Griffith, R Nix, E Bernhardt, and AJ Burgin. 2012. Development and application of a simulation environment (NEO) for integrating empirical and computational investigations of system-level complexity. Proceedings of the International Conference on Information Science and Applications. doi:10.1109/ICISA.2012.6220928.
  59. Lindberg, TT, ES Bernhardt, R Bier, AM Helton, R Merola, A Vengosha, and RT Di Giulioa. 2011. Cumulative impacts of mountaintop mining on an Appalachian watershed. Proceedings of the National Academy of Sciences of the United States of America. 108(52): 20929–20934.
  60. Beaulieu, JJ, JL Tank, SK Hamilton, WM Wollheim, RO Hall Jr., PJ Mulholland, BJ Peterson, LR Ashkenas, LW Cooper, CN Dahm, WK Dodds, NB Grimm, SL Johnson, WH McDowell, GC Poole, HM Valett, CP Arango, MJ Bernot, AJ Burgin, C Crenshaw, AM Helton, L Johnson, JM. O'Brien, JD Potter, RW Sheibley, DJ Sobota, and SM Thomas. 2011. Nitrous oxide emission from denitrification in stream and river networks. Proceedings of the National Academy of Sciences of the United States of America. 108(1): 214-219.
  61. Helton, AM, GC Poole, JL Meyer, WM Wollheim, BJ Peterson, PJ Mulholland, ES Bernhardt, JA Stanford, C Arango, LR Ashkenas, LW Cooper, WK Dodds, SV Gregory, RO Hall Jr, SK Hamilton, SL Johnson, WH McDowell, JD Potter, JL Tank, SM Thomas, HM Valett, JR Webster, and L Zeglin. 2011. Thinking outside the channel: Modeling nitrogen cycling in networked river ecosystems.  Frontiers in Ecology and the Environment. 9(4): 229–238.
  62. Bernot, MJ, DJ Sobota, RO Hall Jr., PJ Mulholland, WK Dodds, JR Webster, JL Tank, LR Ashkenas, LW Cooper, CN Dahm, SV Gregory, NB Grimm, SK Hamilton, SL Johnson, WH McDowell, JL Meyer, BJ Peterson, GC Poole, HM Valett, C Arango, JJ Beaulieu, AJ Burgin, C Crenshaw, AM Helton, L Johnson, J Meriram, BR Niederlehner, JM O’Brien, JD Potter, RW Sheibley, SM Thomas, and K Wilson. 2010. Inter-regional comparison of land-use effects on stream metabolism. Freshwater Biology. 55: 1874-1890.
  63. Small, GE, AM Helton, and C Kazanci. 2009. Can consumer stoichiometric regulation control nutrient spiraling in streams?  Journal of the North American Benthological Society. 28(4):747–765.
  64. Mulholland, PJ, RO Hall, DJ Sobota, WK Dodds, SEG Findlay, NB Grimm, SK Hamilton, WH  McDowell, JM O’Brien, JL Tank, LR Ashkenas, LW Cooper, CN Dahm, SV Gregory, SL Johnson, JL Meyer, BJ Peterson, GC Poole, HM Valett, JR Webster, CP Arango, JJ Beaulieu, MJ Bernot, AJ Burgin, CL Crenshaw, AM Helton, LT Johnson, BR Niederlehner, JD Potter, RW Sheibley, and SM Thomas. 2009. Nitrate removal in stream ecosystems measured by 15N addition experiments: Denitrification. Limnology and Oceanography. 54(3): 666–680.
  65. Hall, RO, JL Tank, DJ Sobota, PJ Mulholland, JM O’Brien, WK Dodds, JR Webster, HM Valett, GC Poole, BJ Peterson, JL Meyer, WH McDowell, SL Johnson, SK Hamilton, NB Grimm, SV Gregory, CN Dahm, LW Cooper, LR Ashkenas, SM Thomas, RW Sheibley, JD Potter, BR Neiderlehner, LT Johnson, AM Helton, CM Crenshaw, AJ Burgin, MJ Bernot, JJ Beaulieu, and CP Arango. 2009. Nitrate removal in stream ecosystems measured by 15N addition experiments: Total uptake. Limnology and Oceanography. 54(3): 653-665.
  66. Mulholland, PJ, AM Helton, GC Poole, RO Hall, Jr., SK Hamilton, BJ Peterson, JL Tank, LR Ashkenas, LW Cooper, CN Dahm, WK Dodds, S Findlay, SV Gregory, NB Grimm, SL Johnson, WH McDowell, JL Meyer, HM Valett, JR Webster, C Arango, JJ Beaulieu, MJ Bernot, AJ Burgin, C Crenshaw, L Johnson, BR Niederlehner, JM O’Brien, JD Potter, RW Sheibley, DJ Sobota, and SM Thomas. 2008. Stream denitrification across biomes and effects of anthropogenic nitrate loading. Nature. 452: 202-206.
  67. Poole, GC, SJ O'Daniel, KL Jones, WW Woessner, ES Bernhardt ES, AM Helton, JA Stanford, BR Boer, TJ Beechie. 2008. Hydrologic Spirals: The Role of Multiple Interactive Flow Paths in Stream Ecosystems. River Research and Applications. 24: 1018-1031.